With the world's population increasing and a large percentage of this population living in urban areas, the need for efficient and low-consumption agricultural methods is becoming more and more necessary. Vertical farming as a solution to the challenges of food security and environmental change, using vertical spaces and controlled environments, by reducing the consumption of various resources, reducing greenhouse gas emissions and environmental pollution of traditional agriculture, while helping the environment, enables the production of agricultural products. Microgreens, which have high amounts of vitamins, minerals and antioxidant compounds, can be used as a new generation of healthy foods and an ideal option for cultivation in vertical systems. Several factors, including the selection of appropriate species, control of the light spectrum (especially blue and red spectra), temperature, humidity, seed density and type of growing medium, affect the growth and quality of microgreens. Along with its advantages, vertical farming faces challenges such as high initial cost, high energy consumption, limited crop yield, gaining consumer trust, and competition with traditional crops. However, technological advancements and further research will help reduce these challenges and expand vertical farming. This article examines the vertical farming system, especially its potential for producing microgreens.
Al-Chalabi, M., 2015. Vertical farming: Skyscraper sustainability?. Sustainable Cities and Society, 18, 74-77.
Al-Kodmany, K., 2018. The Vertical Farm: A Review of Developments and Implications for the Vertical City. Buildings, 8, 24.
Allegaert, S.D., 2020. The Vertical Farm Industry: Exploratory Research of a Wicked Situation. Wageningen University and Research: Wageningen, The Netherlands.
Alrifai, O., Mats, L., Liu, R., Hao, X., Marcone, M.F., Tsao, R., 2021. Effect of combined light-emitting diodes on the accumulation of glucosinolates in Brassica microgreens. Food Production, Processing and Nutrition, 3(1), 1–16.
Ares, G., Ha, B., Jaeger, S.R., 2021. Consumer attitudes to vertical farming (indoor plant factory with artificial lighting) in China, Singapore, UK, and USA: A multi-method study. Food Research International, 150, 110811. https://doi.org/10.1016/j.foodres.2021.110811
Asgari, N., Hayibo, K.S., Groza, J., Rana, S., Pearce, J.M., 2024. Greenhouse Applications of Solar Photovoltaic Driven Heat Pumps in Northern Environments. Western University: London, ON, Canada, to be published.
Astee, L.Y., Kishnani, N.T., 2010. Building integrated agriculture: Utilising rooftops for sustainable food crop cultivation in Singapore. Journal of Green Building, 5(2), 105-113.
Avgoustaki, D.D., Xydis, G., 2020. Indoor Vertical Farming in the Urban Nexus Context: Business Growth and Resource Savings. Sustainability, 12, 1965.
Azizi, S., Lastochkina, O.V., Seyed Hajizadeh, H., Aliniaeifard, S., 2024. Proper quality of LED light to produce high-quality ornamental plants in controlled environment agricultural systems: A review. Greenhouse Plant Production Journal, 1(2), 35–50. https://doi.org/10.61186/gppj.1.2.35
Azizoglu, U., Yilmaz, N., Simsek, O., Ibal, J.C., Tagele, S.B., Shin, J.-H., 2021. The fate of plant growth-promoting rhizobacteria in soilless agriculture: Future perspectives. 3 Biotech, 11, 382.
Baenas, N., Villa˜no, D., García-Viguera, C., Moreno, D.A., 2016. Optimizing elicitation and seed priming to enrich broccoli and radish sprouts in glucosinolates. Food Chemistry, 204, 314–319.
Bailey, R.L., West, K.P., Jr., Black, R.E., 2015. The Epidemiology of Global Micronutrient Deficiencies. Annals of Nutrition and Metabolism, 66 (Suppl. S2), 22–33.
Barbosa, G.L., Almeida Gadelha, F.D., Kublik, N., Proctor, A., Reichelm, L., Weissinger, E., Wohlleb, G.M., Halden, R.U., 2015. Comparison of Land, Water, and Energy Requirements of Lettuce Grown Using Hydroponic vs. Conventional Agricultural Methods. International Journal of Environmental Research and Public Health, 12, 6879–6891.
Beacham, A.M., Vickers, L.H., Monaghan, J.M., 2019. Vertical farming: A summary of approaches to growing skywards. Journal of Horticultural Science and Biotechnology, 94, 277–283.
Benis, K., Ferrao, P., 2018. Commercial farming within the urban built environment e taking stock of an evolving field in northern countries. Global Food Security, 17, 30–37. https://doi.org/10.1016/j.gfs.2018.03.005
Benke, K., Tomkins B., 2017. Future food-production systems: Vertical farming and controlled-environment agriculture. Sustainability: Science, Practice and Policy, 13, 13–26.
Berba, K.J., Uchanski, M.E., 2012. Post-Harvest Physiology of Microgreens. Journal of Young Investigators, 24, 5.
Besthorn, F.H., 2013. Vertical farming: Social work and sustainable urban agriculture in an age of global food crises. Australian Social Work, 66(2), 187-203.
Bhaswant, M., Shanmugam, D.K., Miyazawa, T., Abe, C., Miyazawa, T., 2023. Microgreens—A Comprehensive Review of Bioactive Molecules and Health Benefits. Molecules, 28(2), 867.
Bol, R., Gruau, G., Mellander, P.E., Dupas, R., Bechmann, M., Skarbøvik, E., Bieroza, M., Djodjic, F., Glendell, M., Jordan, P., et al., 2018. Challenges of reducing phosphorus based water eutrophication in the agricultural landscapes of Northwest Europe. Frontiers in Marine Science, 5, 276.
Brazaitytė, A., Sakalauskienė, S., Samuolienė, G., Jankauskienė, J., Viršilė, A., Novičkovas, A., Sirtautas, R., Miliauskienė, J., Vaštakaitė, V., Dabašinskas, L., Duchovskis, P., 2015. The Effects of LED Illumination Spectra and Intensity on Carotenoid Content in Brassicaceae Microgreens. Food Chemistry, 173, 600-606.
Brazaitytė, A., Sakalauskienė, S., Samuolienė, G., Jankauskienė, J., Viršilė, A., Novičkovas, A., Sirtautas, R., Miliauskienė, J., Vaštakaitė, V., Dabašinskas, L., Duchovskis, P., 2015. The Effects of LED Illumination Spectra and Intensity on Carotenoid Content in Brassicaceae Microgreens. Food Chemistry, 173, 600-606. https://doi.org/10.1016/j.foodchem.2014.10.077
Bulgari, R., Negri, M., Santoro, P., Ferrante, A., 2021. Quality Evaluation of Indoor-Grown Microgreens Cultivated on Three Different Substrates. 7(5), 96.
Butturini, M., Marcells, L.F.M., 2020. Vertical farming in Europe: present status and outlook. Elsevier Inc., 77–88.
Carotti, L., et al., 2023. Improving water use efficiency in vertical farming: effects of growing systems, far-red radiation and planting density on lettuce cultivation. Agricultural Water Management, 285, 108365.
Casey, L., Freeman, B., Francis, K., Brychkova, G., McKeown, P., Spillane, C., Bezrukov, A., Zaworotko, M., Styles, D., 2022. Comparative environmental footprints of lettuce supplied by hydroponic controlled-environment agriculture and field-based supply chains. Journal of Cleaner Production, 369, 133214. https://doi.org/10.1016/j.jclepro.2022.133214
Chaudhry, A.R., Mishra, V.P., 2019. A comparative analysis of vertical agriculture systems in residential apartments. In: Proceedings of the 2019 Advances in Science and Engineering Technology International Conferences (ASET), Dubai, United Arab Emirates, 26 March–10 April 2019.
Chen, P., Zhu, G., Kim, H.-J., Brown, P.B., Huang, J.-Y., 2020. Comparative life cycle assessment of aquaponics and hydroponics in the midwestern United States. Journal of Cleaner Production, 275, 122888. https://doi.org/10.1016/j.jclepro.2020.122888
Chowdhury, M.E.H., et al., 2020. Design, construction and testing of IoT based automated indoor vertical hydroponics farming test-bed in Qatar. Sensors, 20(19), 5637. https://doi.org/10.3390/s20195637
Ciuta, F., Arghir, L.D., Tudor, C.A., Lagunovschi-Luchian, V., 2020. Research on microgreens farming in vertical hydroponic system. Journal of Horticulture, Forestry and Biotechnology.
Corvalan, C., Hales, S., McMichael, A.J., 2005. Ecosystems and Human Well-Being: Health Synthesis. World Health Organization: Geneva, Switzerland.
Delorme, M., Santini, A., 2022. Energy-efficient automated vertical farms. Omega, 109, 102611.
Den Besten, J., 2019. Vertical farming development; the Dutch approach. In: Anpo, M., Hirokazu, F., Teruo, W. (Eds.), Plant Factory Using Artificial Light. Elsevier, pp. 307–317. https://doi.org/10.1016/B978-0-12-13973-8.00027-0
Despommier, D., 2010. The Vertical Farm: Feeding the World in the 21st Century. Thomas Dunne Books: New York, NY.
Di Gioia, F., De Bellis, P., Mininni, C., Santamaria, P., Serio, F., 2017. Physicochemical, agronomical and microbiological evaluation of alternative growing media for the production of rapini (Brassica rapa L.) microgreens. Journal of the Science of Food and Agriculture, 97, 1212–1219. https://doi.org/10.1002/jsfa.7852
Ebert, A.W., 2022. Sprouts and Microgreens—Novel Food Sources for Healthy Diets. Plants, 11, 571.
European Commission, 2008. Commission Regulation (EC) No 889/2008 of 5 September 2008 laying down detailed rules for the implementation of Council Regulation (EC) No 834/2007 on organic production and labelling of organic products with regard to organic production, labelling and control. Official Journal of the European Union.
Ferrón-Carrillo, F., Guil-Guerrero, J.L., González-Fernández, M.J., Lyashenko, S., Battafarano, F., da Cunha-Chiamolera, T.P.L., Urrestarazu, M., 2021. LED enhances plant performance and both carotenoids and nitrates profiles in lettuce. Plant Foods for Human Nutrition, 1–9.
Flores, M., Urrestarazu, M., Amorós, A., Escalona, V., 2022. High intensity and red enriched LED lights increased the growth of lettuce and endive. Italian Journal of Agronomy, 17, 1915. https://doi.org/10.4081/ija.2022.1915
Galieni, A., Falcinelli, B., Stagnari, F., Datti, A., Benincasa, P., 2020. Sprouts and microgreens: Trends, opportunities, and horizons for novel research. Agronomy, 10, 1424.
Gan, C.I., Soukoutou, R., Conroy, D.M., 2023. Sustainability framing of controlled environment agriculture and consumer perceptions: a review. Sustainability, 15, 304.
Gerovac, J.R., Craver, J.K., Boldt, J.K., Lopez, R.G., 2016. Light intensity and quality from sole-source light-emitting diodes impact growth, morphology, and nutrient content of Brassica microgreens. HortScience, 51, 497–503.
Gertphol, S., Chulaka, P., Changmai, T., 2018. Predictive models for lettuce quality from Internet of Things-based hydroponic farm. In: 2018 22nd International Computer Science and Engineering Conference (ICSEC). pp. 1–5. https://doi.org/10.1109/ICSEC.2018.8712676.
Goodman, W., Minner, J., 2019. Will the urban agricultural revolution be vertical and soilless? A case study of controlled environment agriculture in New York City. Land Use Policy, 83, 160–173.
Graamans, L., Baeza, E., van den Dobbelsteen, A., Tsafaras, I., Stanghellini, C., 2018. Plant factories versus greenhouses: comparison of resource use efficiency. Agricultural Systems, 160, 31–43.
Gruda, N., 2019. Increasing sustainability of growing media constituents and stand-alone substrates in soilless culture systems. Agronomy, 9, 298.
Grunert, O., Reheul, D., Van Labeke, M.-C., Perneel, M., Hernandez-Sanabria, E., Vlaeminck, S.E., Boon, N., 2016. Growing media constituents determine the microbial nitrogen conversions in organic growing media for horticulture. Microbial Biotechnology, 9, 389–399.
Hallikainen, E., 2018. Life Cycle Assessment on Vertical Farming. Aalto University: Aalto, Finland.
Hatfield, J.L., Prueger, J.H., 2015. Temperature extremes: Effect on plant growth and development. Weather and Climate Extremes, 10, 4–10.
Healy, R.G., Rosenberg, J.S., 2013. Land Use and the States. Routledge: New York, NY.
Heo, J.-W., Kang, D.-H., Bang, H.-S., Hong, S.-G., Chun, C.-H., Kang, K.-K., 2012. Early growth, pigmentation, protein content, and phenylalanine ammonia-lyase activity of red curled lettuces grown under different lighting conditions. Korean Journal of Horticultural Science and Technology, 30, 6–12. https://doi.org/10.7235/hort.2012.11118
Hoang, G.M., Vu, T.T., 2022. Selection of suitable growing substrates and quality assessment of andBrassicaand microgreens cultivated in greenhouse. Academia Journal of Biology, 44, 133–142.
Huang, H., Jiang, X., Xiao, Z., Yu, L., Pham, Q., Sun, J., Chen, P., Yokoyama, W., Yu, L.L., Luo, Y.S., Wang, T.T., 2016. Red cabbage microgreens lower circulating low-density lipoprotein (LDL), liver cholesterol, and inflammatory cytokines in mice fed a high-fat diet. Journal of Agricultural and Food Chemistry, 64(48), 9161–9171. https://doi.org/10.1021/acs.jafc.6b03805
Hydroponics Factory, 2024. Hydroponic systems and solutions. Available at: https://www.hydroponicsfactory.com
iFarm, 2024. Vertical farming technology. Available at: https://www.ifarm.fi
Johnson, G., 2023. Edible Garden approves reverse stock split. Produce Blue Book, Carol Stream, IL, USA.
Jones-Baumgardt, C., Llewellyn, D., Ying, Q., Zheng, Y., 2019. Intensity of sole-source light-emitting diodes affects growth, yield, and quality of Brassicaceae microgreens. HortScience, 54, 1168–1174. https://doi.org/10.21273/HORTSCI13788-18
Jowell, A., Zhou, B., Barry, M., 2017. The impact of megacities on health: Preparing for a resilient future. The Lancet Planetary Health, 1, e176–e178. https://doi.org/10.1016/S2542-5196(17)30080-3
Kabir, M.S.N., Reza, M.N., Chowdhury, M., Ali, M., Samsuzzaman, Ali, M.R., Lee, K.Y., Chung, S.O., 2023. Technological trends and engineering issues on vertical farms: A review. Horticulturae, 9, 1229.
Kalantari, F., Tahir, O.M., Lahijani, A.M., Kalantari, S., 2017. A review of vertical farming technology: A guide for implementation of building integrated agriculture in cities. Advanced Engineering Forum, 24, 76–91.
Katz, R., Bradley, J., 2013. The Metropolitan Revolution: How Cities and Metropolitan Areas Are Fixing Broken Politics and Fragile Economy. Brookings Institution Press: Washington, DC.
Kikuchi, Y., Kanematsu, Y., Yoshikawa, N., Okubo, T., Takagaki, M., 2018. Environmental and resource use analysis of plant factories with energy technology options: A case study in Japan. Journal of Cleaner Production, 186, 703–717.
Kim, H.-S., Kim, K.-R., Lim, G.-H., Kim, J.-W., Kim, K.-H., 2015. Influence of airborne dust on the metal concentrations in crop plants cultivated in a rooftop garden in Seoul. Soil Science and Plant Nutrition, 61, 88–97.
Kim, M.J., Mikš-Krajnik, M., Kumar, A., Yuk, H.G., 2016. Inactivation by 405 ± 5 nm light emitting diode on andEscherichia coliand O157:H7, andSalmonella Typhimuriumand, and andShigella sonneiand under refrigerated condition might be due to the loss of membrane integrity. Food Control, 59, 99–107.
Kim, S.-J., Hahn, E.-J., Heo, J.-W., Paek, K.-Y., 2004. Effects of LEDs on net photosynthetic rate, growth and leaf stomata of chrysanthemum plantlets andin vitroand. Scientia Horticulturae, 101, 143–151. https://doi.org/10.1016/j.scienta.2003.10.003
Kou, L., Yang, T., Luo, Y., Liu, X., Huang, L., Codling, E., 2014. Pre-harvest calcium application increases biomass and delays senescence of broccoli microgreens. Postharvest Biology and Technology, 87, 70–78.
Kozai, T., Niu, G., Takagaki, M. (Eds.), 2019. andPlant Factory: An Indoor Vertical Farming System for Efficient Quality Food Productionand. Academic Press: Cambridge, MA, USA.
Kozai, T., Niu, G., Takagaki, M., 2016. andPlant Factory: An Indoor Vertical Farming System for Efficient Quality Food Productionand. Academic Press: San Diego, CA, USA.
Kulak, M., Graves, A., Chatterton, J., 2013. Reducing greenhouse gas emissions with urban agriculture: A Life Cycle Assessment perspective. Landscape and Urban Planning, 111, 68–78.
Kusuma, P., Pattison, P.M., Bugbee, B., 2020. From physics to fixtures to food: Current and potential LED efficacy. Horticulture Research, 7, 56.
Kyriacou, M.C., Rouphael, Y., Di Gioia, F., Kyratzis, A., Serio, F., Renna, M., De Pascale, S., Santamaria, P., 2016. Micro-scale vegetable production and the rise of microgreens. Trends in Food Science and Technology, 57, 103–115.
Lada, R.R., Maijers, W., Nieboer, S., 2018. Innovative Indoor Horticultural Systems (IHORT) for the 21st Century. Current Investigations in Agriculture and Current Research, 4, 576–581. https://doi.org/10.32474/CIACR.2018.04.000194
Le, T.N., Chiu, C.H., Hsieh, P.C., 2020. Bioactive compounds and bioactivities of andBrassica oleraceaand L. var. anditalicaand sprouts and microgreens: An updated overview from a nutraceutical perspective. Plants, 9, 946.
Lee, K.O., Mai, K.M., Park, S., 2023. Green space accessibility helps buffer declined mental health during the COVID-19 pandemic: Evidence from big data in the United Kingdom. andNature Mental Healthand, 1, 124–134.
Lee-Kwan, S.H., Moore, L.V., Blanck, H.M., Harris, D.M., Galuska, D., 2017. Disparities in state-specific adult fruit and vegetable consumption—United States, 2015. andMorbidity and Mortality Weekly Report (MMWR)and, 66, 1241–1247.
Liu, T., Yang, M., Han, Z., Ow, D.W., 2016. Rooftop production of leafy vegetables can be profitable and less contaminated than farm-grown vegetables. andAgronomy for Sustainable Developmentand, 36, 41.
Lobiuc, A., Vasilache, V., Oroian, M., Stoleru, T., Burducea, M., Pintilie, O., Zamfirache, M.-M., 2017. Blue and red LED illumination improves growth and bioactive compounds contents in acyanic and cyanic andOcimum basilicumand L. microgreens. andMoleculesand, 22, 2111. https://doi.org/10.3390/molecules22122111
Martin, M., Molin, E., 2019. Environmental assessment of an urban vertical hydroponic farming system in Sweden. andSustainabilityand, 11, 4124.
McGehee, C.S., Raudales, R.E., Elmer, W.H., McAvoy, R.J., 2019. Efficacy of biofungicides against root rot and damping-off of microgreens caused by andPythiumand spp. andCrop Protectionand, 121, 96–102.
Mousavi, S.N., Alamzadeh Ansari, N., 2025. Investigating the effect of density and cultivar on onion seedling production in a floating culture system. andGreenhouse Plant Production Journaland, 2(1), 1–17. https://doi.org/10.61186/gppi.2.1.1
Muchjajib, U., Muchjajib, S., Suknikom, S., Butsai, J., 2015. Evaluation of organic media alternatives for the production of microgreens in Thailand. andActa Horticulturaeand, 1102, 157–162. https://doi.org/10.17660/ActaHortic.2015.1102.19
Mukherji, N., Morales, A., 2010. andZoning for Urban Agricultureand. Zoning Practice 3. American Planning Association: Chicago, IL.
Muller, A., Ferré, M., Engel, S., Gattinger, A., Holzkämper, A., Huber, R., Müller, M., Six, J., 2017. Can soil-less crop production be a sustainable option for soil conservation and future agriculture? andLand Use Policyand, 69, 102–105.
Muneer, S., Kim, E., Park, J., Lee, J., 2014. Influence of green, red and blue light emitting diodes on multiprotein complex proteins and photosynthetic activity under different light intensities in lettuce leaves (Lactuca sativaand L.). andInternational Journal of Molecular Sciencesand, 15, 4657–4670. https://doi.org/10.3390/ijms15034657
Murphy, C.J., Pill, W.G., 2010. Cultural practices to speed the growth of microgreen arugula (roquette; andEruca vesicariaand subsp andsativaand). Journal of Horticultural Science and Biotechnology, 85, 171–176.
Naikoo, N.B., Kanth, R.H., Bahar, F.A., Bhat, M.A., Nazir, A., Mahdi, S.S., Amin, Z., Singh, L., Raja, W., et al., 2022. Vertical farming: The future of agriculture: A review. andPharma Innovation Journaland, 11, 1175–1195.
Nicola, S., Pignata, G., Ferrante, A., Bulgari, R., Cocetta, G., Ertani, A., 2020. Water use efficiency in greenhouse systems and its application in horticulture. andAgroLife Scientific Journaland, 9, 248–262.
Nils, E., 2018. Groente kweken in voedselflats: kansrijk, maar kostbaar. andTrouwand. Available at: https://www.trouw.nl/groen/groente-kweken-in-voedselflats-kansrijk-maar-kostbaarwa591e239
Nishio, J.N., 2000. Why are higher plants green? Evolution of the higher plant photosynthetic pigment complement. andPlant, Cell and Environmentand, 23, 539–548. https://doi.org/10.1046/j.1365-3040.2000.00563.x
Noichinda, S., Bodhipadma, K., Mahamontri, C., Narongruk, T., Ketsa, S., 2007. Light during storage prevents loss of ascorbic acid, and increases glucose and fructose levels in Chinese kale (andBrassica oleraceaand var. andalboglabraand). andPostharvest Biology and Technologyand, 44, 312–315.
Norman, K., Haß, U., Pirlich, M., 2021. Malnutrition in older adults—recent advances and remaining challenges. andNutrientsand, 13, 2764.
Oh, M.M., Carey, E.E., Rajashekar, C.B., 2010. Regulated water deficits improve phytochemical concentration in lettuce. andJournal of the American Society for Horticultural Scienceand, 135, 223–229.
Pattison, P.M., Tsao, J.Y., Brainard, G.C., Bugbee, B., 2018. LEDs for photons, physiology and food. andNatureand, 563, 493–500. https://doi.org/10.1038/s41586-018-0706-x
Palmitessa, O.D., Renna, M., Crupi, P., Lovece, A., Corbo, F., Santamaria, P., 2020. Yield and quality characteristics of andBrassicaand microgreens as affected by the NH4:NO3 molar ratio and strength of the nutrient solution. andFoodsand, 9, 677.
Pannico, A., Graziani, G., El-Nakhel, C., Giordano, M., Ritieni, A., Kyriacou, M., Rouphael, Y., 2020. Nutritional stress suppresses nitrate content and positively impacts ascorbic acid concentration and phenolic acids profile of lettuce microgreens. andItalus Hortusand, 27, 41–52.
Partap, M., Sharma, D., HN, D., Thakur, M., Verma, V., Ujala, Bhargava, B., 2023. Microgreen: A tiny plant with superfood potential. andJournal of Functional Foodsand, 107, 105697.
Pinstrup-Andersen, P., 2018. Is it time to take vertical indoor farming seriously? andGlobal Food Securityand, 17, 233–235.
Pinto, E., Almeida, A.A., Aguiar, A.A., Ferreira, I.M.P.L.V.O., 2015. Comparison between the mineral profile and nitrate content of microgreens and mature lettuces. Journal of Food Composition and Analysis 37, 38–43. <https://doi.org/10.1016/j.jfca.2014.06.018
Poulet, L., Massa, G.D., Morrow, R.C., Bourget, C.M., Wheeler, R.M., Mitchell, C.A., 2014. Significant reduction in energy for plant-growth lighting in space using targeted LED lighting and spectral manipulation. Life Sciences in Space Research 2, 43–53. <https://doi.org/10.1016/j.lssr.2014.06.002>
Rabobank, 2018. Vertical Farming in the Netherlands. Rabobank, Venlo, the Netherlands. Lecture presented on 27 June 2018.
Rajan, P., Lada, R.R., MacDonald, M.T., 2019. Advancement in indoor vertical farming for microgreen production. American Journal of Plant Sciences 10, 1397–1408. <https://doi.org/10.4236/ajps.2019.108100>
Renna, M., Paradiso, V.M., 2020. Ongoing research on microgreens: Nutritional properties, shelf-life, sustainable production, innovative growing and processing approaches. Foods 9, 826.
Riggio, G., Jones, S., Gibson, K., 2019. Risk of human pathogen internalization in leafy vegetables during lab-scale hydroponic cultivation. Horticulturae 5, 25.
Righini, C., Stanghellini, S., Hemming, L., Graamans, L.F.M., Marcelis, 2023. Resources for plant-based food: Estimating resource use to meet the requirements of urban and peri-urban diets. Food and Energy Security 12, e462. <https://doi.org/10.1002/fes3.462>
Romeo, D., Vea, E.B., Thomsen, M., 2018. Environmental impacts of urban hydroponics in Europe: A case study in Lyon. Procedia CIRP 69, 540–545.
Roosta, H.R., Sharifinejad, S., Azizi Ilami, M.R., Bikdeloo, M., 2024. Different ratios of vermicompost in planting substrates affect the growth and physiological characteristics of Spathiphyllum (Spathiphyllum wallisii). Greenhouse Plant Production Journal 1(4), 49–72.
Rusu, T., Cowden, R.J., Moraru, P.I., Maxim, M.A., Ghaley, B.B., 2021. Overview of multiple applications of basil species and cultivars and the effects of production environmental parameters on yields and secondary metabolites in hydroponic systems. Sustainability 13, 11332.
Saini, R.K., Keum, Y.S., 2018. Carotenoid extraction methods: A review of recent developments. Food Chemistry 240, 90–103.
Sambo, P., Nicoletto, C., Giro, A., Pii, Y., Valentinuzzi, F., Mimmo, T., Lugli, P., Orzes, G., Mazzetto, F., Astolfi, S., et al., 2019. Hydroponic solutions for soilless production systems: Issues and opportunities in a smart agriculture perspective. Frontiers in Plant Science 10, 923.
Samuolien˙ e, G., Brazaityt˙ e, A., Viršil˙ e, A., Miliauskien˙ e, J., Vaštakait˙ e-Kairien˙ e, V., Duchovskis, P., 2019. Nutrient levels in Brassicaceae microgreens increase under tailored light-emitting diode spectra. Frontiers in Plant Science 10, 1475.
Samuoliene, G., Virsile, A., Haimi, P., Miliauskiene, J., 2020. Photoresponse to different lighting strategies during red leaf lettuce growth. Journal of Photochemistry and Photobiology B: Biology 202, 111726.
Sandoval-Insausti, H., Chiu, Y.-H., Lee, D.H., Wang, S., Hart, J.E., Mínguez-Alarcón, L., Laden, F., Korat, A.V.A., Birmann, B., Eliassen, A.H., et al., 2021. Intake of fruits and vegetables by pesticide residue status in relation to cancer risk. Environment International 156, 106744.
Savvas, D., 2003. Hydroponics: A modern technology supporting the application of integrated crop management in greenhouse. Journal of Food, Agriculture and Environment 1, 80–86.
SharathKumar, M., Heuvelink, E., Marcelis, L.F.M., 2020. Vertical farming: Moving from genetic to environmental modification. Trends in Plant Science 25, 724–727.
Sijmonsma, A., 2018. Vertical farming is difficult in the Netherlands. Hortidaily. Available at: <https://www.hortidaily.com/article/44518/Vertical-farming-is-difficult-in-the-Netherlands
Singh, D., Basu, C., Meinhardt-Wollweber, M., Roth, B., 2015. LEDs for energy efficient greenhouse lighting. Renewable and Sustainable Energy Reviews 49, 139–147. <https://doi.org/10.1016/j.rser.2015.04.117
Smith, H.L., et al., 2017. Don’t ignore the green light: Exploring diverse roles in plant processes. Journal of Experimental Botany 68, 2099–2110.
Soufi, H.R., Ali, A., Shojaee, M., 2024. Improving the growth and photosynthesis characteristics of lettuce cultivars in the aeroponic system: With emphasis on the use of supplementary LED lights and replacement methods of the nutrient solution. Greenhouse Plant Production Journal 1(3), 28–44.
Specht, K., et al., 2019. How will we eat and produce in the cities of the future? From edible insects to vertical farming—a study on the perception and acceptability of new approaches. Sustainability 11, 4315.
Stanghellini, C., Van ’t Ooster, B., Heuvelink, E., 2019. Vertical farms. In: Greenhouse Horticulture. Wageningen Academic Publishers, The Netherlands. <https://doi.org/10.3920/978-90-8686-879-7>
Sun, J., Xiao, Z., Lin, L.-Z., Lester, G.E., Wang, Q., Harnly, J.M., Chen, P., 2013. Profiling polyphenols in five brassica species microgreens by UHPLC-PDA-ESI/HRMSn. Journal of Agricultural and Food Chemistry 61, 10960–10970.
Szpyrka, E., Kurdziel, A., Rupar, J., Slowik-Borowiec, M., 2015. Pesticide residues in fruit and vegetable crops from the central and eastern region of Poland. Roczniki Panstwowego Zakladu Higieny 66, 107–113.
Tan, L., Nuffer, H., Feng, J., Kwan, S.H., Chen, H., Tong, X., Kong, L., 2020. Antioxidant properties and sensory evaluation of microgreens from commercial and local farms. Food Science and Human Wellness 9(1), 45–51.
Tavan, M., Wee, B., Brodie, G., Fuentes, S., Pang, A., Gupta, D., 2021. Optimizing sensor-based irrigation management in a soilless vertical farm for growing microgreens. Frontiers in Sustainable Food Systems 4, 622720.
The United Nations, 2017. World Population Prospects: The 2017 Revision. United Nations, New York.
Tolentino, L.K.S., et al., 2021. Hylo: Implementation of LoRaWAN in an automated hydroponics system. In: 2021 IEEE International Conference on Mobile Networks and Wireless. https://doi.org/10.1109/ICMNWC52512.2021.9688540>
Touliatos, D., Dodd, I.C., McAinsh, M., 2016. Vertical farming increases lettuce yield per unit area compared to conventional horizontal hydroponics. Food and Energy Security 5(3), 184–191.
Treadwell, D., Hochmuth, R., Landrum, L., Laughlin, W., 2020. Microgreens: A new specialty crop: HS1164, rev. 9/2020. EDIS 2020(1), 1–3.
Treadwell, D.D., Hochmuth, R., Landrum, L., Laughlin, W., 2010. Microgreens: A new specialty crop. University of Florida IFAS Extension HS1164.3.
Tuomisto, H.L., 2019. Vertical farming and cultured meat: Immature technologies for urgent problems. One Earth 1, 275–277.
van Delden, S.H., et al., 2021. Current status and future challenges in implementing and upscaling vertical farming systems. Nature Food 2, 944–956.
Van Gerrewey, T., Boon, N., Geelen, D., 2022. Vertical farming: The only way is up? Agronomy 12(1). <https://doi.org/10.3390/agronomy12010002>
Vatistas, C., Avgoustaki, D.D., Bartzanas, T., 2022. A systematic literature review on controlled-environment agriculture: How vertical farms and greenhouses can influence the sustainability and footprint of urban microclimate with local food production. Atmosphere 13(8), 1258. <https://doi.org/10.3390/atmos13081258>
Virsile, A., Sirtautas, R., 2013. Light irradiance level for optimal growth and nutrient contents in borage microgreens. Proceedings of the International Scientific Conference: Rural Development 6, 272.
Wang, X., Onychko, V., Zubko, V., Wu, Z., Zhao, M., 2023. Sustainable production systems of urban agriculture in the future: A case study on the investigation and development countermeasures of the plant factory and vertical farm in China. Frontiers in Sustainable Food Systems 7, 16.
Weber, C.F., 2016. Nutrient content of cabbage and lettuce microgreens grown on vermicompost and hydroponic growing pads. Journal of Horticulture 3, 1–5. <https://doi.org/10.4172/2376-0354.1000190>
Wells, J.C., Sawaya, A.L., Wibaek, R., Mwangome, M., Poullas, M.S., Yajnik, C.S., Demaio, A., 2019. The double burden of malnutrition: Aetiological pathways and consequences for health. The Lancet 395, 75–88.
Wildeman, R., 2020. Vertical Farming: A Future Perspective or a Mere Conceptual Idea? University of Twente, Enschede, The Netherlands.
Willett, W., et al., 2019. Food in the anthropocene: The EAT–Lancet Commission on healthy diets from sustainable food systems. The Lancet 393, 447–492.
Xiao, Z., Lester, G.E., Luo, Y., Wang, Q., 2012. Assessment of vitamin and carotenoid concentrations of emerging food products: Edible microgreens. Journal of Agricultural and Food Chemistry 60, 7644–7651. <https://doi.org/10.1021/jf300459b>
Xiao, Z., Nou, X., Luo, Y., Wang, Q., 2014a. Comparison of the growth of Escherichia coli O157:H7 and O104:H4 during sprouting and microgreen production from contaminated seeds.
Xydis, G., Strasszer, D., Avgoustaki, D.D., Nanaki, E., 2021. Mass deployment of plant factories as a source of load flexibility in the grid under an energy-food nexus. A technoeconomics-based comparison. Sustainable Energy Technologies and Assessments 47, 101431.
Yan, H., Li, W., Chen, H., Liao, Q., Xia, M., Wu, D., Liu, C., Chen, J., Zou, L., Peng, L., et al., 2022. Effects of storage temperature, packaging material and wash treatment on quality and shelf life of Tartary buckwheat microgreens. Foods 11, 3630.
Yang, T., Kim, H.-J., 2020. Characterizing nutrient composition and concentration in tomato-, basil-, and lettuce-based aquaponic and hydroponic systems. Water 12, 1259.
Zhang, X., Bian, Z., Li, S., Chen, X., Lu, C., 2019. Comparative analysis of phenolic compound profiles, antioxidant capacities, and expressions of phenolic biosynthesis-related genes in soybean microgreens grown under different light spectra. Journal of Agricultural and Food Chemistry 67, 13577–13588. <https://doi.org/10.1021/acs.jafc.9b05594>
Zhang, X., Bian, Z., Yuan, X., Chen, X., Lu, C., 2020. A review on the effects of light-emitting diode (LED) light on the nutrients of sprouts and microgreens. Trends in Food Science and Technology 99, 203–216.
nooriyan,S. and Roosta,H. R. (2025). Vertical farming with an emphasis in microgreens: A review. Greenhouse Plant Production Journal, 2(1), 73-95. doi: 10.61186/gppj.2.1.73
MLA
nooriyan,S. , and Roosta,H. R. . "Vertical farming with an emphasis in microgreens: A review", Greenhouse Plant Production Journal, 2, 1, 2025, 73-95. doi: 10.61186/gppj.2.1.73
HARVARD
nooriyan S., Roosta H. R. (2025). 'Vertical farming with an emphasis in microgreens: A review', Greenhouse Plant Production Journal, 2(1), pp. 73-95. doi: 10.61186/gppj.2.1.73
CHICAGO
S. nooriyan and H. R. Roosta, "Vertical farming with an emphasis in microgreens: A review," Greenhouse Plant Production Journal, 2 1 (2025): 73-95, doi: 10.61186/gppj.2.1.73
VANCOUVER
nooriyan S., Roosta H. R. Vertical farming with an emphasis in microgreens: A review. JGPP, 2025; 2(1): 73-95. doi: 10.61186/gppj.2.1.73